ACKNOWLEDGEMENTS We would like to thank the staff of participating clinics for their help in collecting biosamples and clinical information. This work was supported by the Russian Science Foundation [Project No 17-75-20223]. The work of Dr Tsepilov was supported by the Russian Ministry of Science and Education under the 5-100 Excellence Programme. We express our gratitude to the members of the Gene ATLAS project (http://geneatlas.roslin.ed.ac.uk/) and personally thank the Albert Tenesa's group, Oriol Canela-Xandri, and Konrad Rawlik for sharing UK Biobank GWAS summary statistics that we have used in our study. We gratefully thank UK Biobank for establishing a powerful resource for genetic and epidemiological studies. The authors wish to correct the information in the Acknowledgement section in their paper.